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Originally Posted by LionHarted
Could it be that FSA is overwriting the function of the Moon Pearl in ALttP, and giving it the power to open portals to the Dark World? It seems that the Four Sword, with its power to pierce darkness, is also connected to the Dark World. Since you don't exactly "use" the Moon Pearl in ALttP until after you get the Master Sword, it would be easy for Nintendo to overwrite the function of the Moon Pearl to allow it to be more connected to portals, and grant the darkness-smiting swords the power to protect the bearers from the Dark World's transformation power.
This would explain why Link doesn't transform in the Dark World in FSA (because he has the Four Sword), and it may be used as a gameplay element in TP to allow Link to wander areas of the Twilight without transforming.
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ALTTP's Moon Pearl actually implied a connection with the "Legendary Hero" like the Magic Mirror. I would be willing to speculate that they are not the same items, even though they have the same names. That said, the exact form in which the Dark World spreads and it's ability to transform people has varied enormously between different games and I can't see any real logic to what causes this variation, other than gameplay of course.